The way I understand it, if a 'front' tire is mounted on the rear, you run it opposite rotation. Reason being, has to do with the design of the plys. (plies?) The front tire is designed mainly for braking forces, whereas the rear tire is designed for acceleration. Two opposite forces. Sounds reasonable to me ...
I would imagine our Enfields would be very forgiving in that, considering the moderate power output and somewhat meager braking capabilities.